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Arithmetic is one of the foundations of our educational systems, but what exactly is it? Numbers are everywhere in our modern societies, but what is our knowledge of numbers really about? This book provides a philosophical account of arithmetical knowledge that is based on the state-of-the-art empirical studies of numerical cognition. It explains how humans have developed arithmetic from humble origins to its modern status as an almost universally possessed knowledge and skill. Central to the account is the realisation that, while arithmetic is a human creation, the development of arithmetic is constrained by our evolutionarily developed cognitive architecture. Arithmetic is a sophisticated cultural development, but it is ultimately based on abilities with numerosities that we already possess as infants and share with many non-human animals. Therefore, arithmetic is not purely conventional, an arbitrary game akin to chess. Instead, arithmetic is deeply connected to our basic cognitive capacities.